This article explores patterns of mediatized participation of European citizens and the way they differ across different media systems, in a multilevel, cross-national comparative research design. Mediatized participation is operationalized as audience practices on the Internet. The media system is conceptualized through the theoretical model of digital mediascapes, which applied to 22 European Union countries produced three clusters/media systems. The audience data are from representative online surveys in 8 eastern and western European countries (N = 9532) collected by the authors and their research partners. Factor and cluster analyses were performed showing types and patterns of mediatized participation. Hierarchical multiple regression analysis and ANOVA were performed to relate the individual level variables to the macro-level clusters of digital media systems. The article shows audiences in the more mediatized, Western cluster are more engaged in participatory practices in comparison to audiences in the Eastern/Southern cluster of European countries which show more extensive information consumption practices.